Cenarchaeum is a monotypic genus of archaeans in the family Cenarchaeaceae. The marine archaean Cenarchaeum symbiosum is psychrophilic and is found inhabiting marine sponges. Cenarchaeum symbiosum was initially detected as a major symbiotic microorganism living within (it is an endosymbiont of) the sponge Axinella mexicana. It has been ubiquitously detected in the world oceans at lower abundances, while in some genera of marine sponges it is one of the most abundant microbiome members. Its genome sequence and diversity has been investigated in detail finding unique metabolic products and its role in ammonia-oxidizing activities.

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