Cosmospora is a genus of ascomycete fungi in the family Nectriaceae. The genus, as circumscribed by Rossman et al. (1998), included all the nectrioid species with small, reddish, non-ornamented sexual fruiting bodies that collapse laterally when dry. However, the genus was shown to be polyphyletic, and the majority of species were re-classified into revived or recently established genera that are monophyletic. Cosmospora sensu Rossman housed members of the following genera: Chaetopsina, Cylindrocladiella, Fusicolla, Macroconia, Mariannaea, Microcera, Pseudocosmospora, Stylonectria, and Volutella. Cosmospora was restricted to species having acremonium-like asexual morphs that grow on polypores and xylariaceous fungi by Gräfenhan in 2011. About 20 species are accepted in the genus (Gräfenhan et al. 2011; Herrera et al. 2015; Zeng and Zhuang et al. 2016; Luo et al. 2019; Lechat et al. 2021).

The name Cosmospora comes from Greek kosmos + spora, meaning ornamented spores.

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