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Parent Taxon

• N/A

Sibling Taxa

• Ateuchini
• Coprini
• Deltochilini
• Epilissini
• Epirinini
• Eucraniini
• Eurysternini
• Gymnopleurini
• Haroldius
• Oniticellini
• Onitini
• Onthophagini
• Onychothecus
• Parachoriini
• Parateuchus
• Phanaeini
• Scarabaeini
• Sisyphini

Child Taxa

• Escarabaeus
• Kheper
• Mentophilus
• Mnematidium
• Mnematium
• Pachylomera
• Pachysoma
• Scarabaeus
• Sceliages

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